Contributors



 

BONAP's contributors

Taxonomic group specialists

Over the past four decades, the following individuals provided valuable contributions. Although, nearly 50% of the original contributors to my 1980 Checklist are now deceased or retired, I continue to honor them here by listing their names and associated areas of specialization. They, along with many others have improved BONAP's efforts and assisted in building our international plant database.

Repositories visited by BONAP or from which plant materials and data have been exchanged

The following herbaria, museums, special collections and other repositories of vascular plants have been visited, consulted, or have provided valuable county-level records for our National Plant Atlas. 

Field botanists

The invaluable contributions of the following 384 individuals cannot be overstated. Tens of thousands of new county-level records have been added to the Atlas because of their collections. Equally important, thousands of erroneous reports have been eliminated. Although I have not always agreed with their conclusions, without their contributions, this Atlas would be hopelessly incomplete and far less accurate. I thank them all.

Photographic contributors

In the early 2000's, I purchased a Nikon slide scanner and initiated a scanning campaign that has spanned ten years. Although the original scanner has been replaced twice, nearly 200,000 slides have been scanned. The vast majority of these slides were contributed generously by hundreds of individuals. BONAP's current image collection contains nearly 300,000 images. Approximately, 100,000 of these are illustrated on our current websites, with the remaining 200,000 to be added later this year. Our goal is to provide quality images for all North American plant taxa as part of our North American Digital Flora and accompanying Floristic Synthesis software. We are extremely grateful to all who have willingly provided their images. Each contributor is credited here for his or her contributions.

WE WELCOME YOUR HELP. When assembling an image gallery this large, it is very easy to misidentify a plant or incorrectly link a scientific name to an image. Therefore, we welcome greatly any corrections. We shall soon include a list of plants for which we have no images or for which we need replacement images. If you would be willing to contribute photos, we would welcome them greatly. Please do not hesitate to contact us.



Taxonomic group specialists

PTERIDOPHYTA CONVOLVULACEAE OXALIDACEAE
Barrington, David S. Austin, Daniel F.
Eiten, George
Mickel, John T. Chao, Jew-Ming
Denton, Melinda F.
Smith, Alan R.
  Lourteig, Alicia
Wagner, Warren H. Jr. CORNACEAE  
Windham, Michael D.
Eyde, Richard H PANDANACEAE
    Stone, Benjamin C.
ADIANTACEAE CRASSULACEAE  
Tryon, Rolla M.
Beauchamp, R. Mitchel
PAPAVERACEAE
(Notholaena) Clausen, Robert T.
Kiger, Robert W.
  Denton, Melinda F.
(Papaver)
CYATHEACEAE Moran, Reid Ownbey, Gerald B.
Tryon, Rolla M.
  (Argemone)
  CROSSOSOMATACEAE  
ISOETACEAE DeBuhr, Larry E.
PASSIFLORACEAE
Taylor, W. Carl
Prigge, Barry A.
Goldman, Douglas H.
     
LYCOPODIACEAE CUCURBITACEAE PEDALIACEAE
Beitel, Joseph M.
Jeffrey, C.
Bretting, Peter K.
  Whitaker, Thomas W.
Hevly, Richard H.
PARKERIACEAE    
Lloyd, Robert M.
CYPERACEAE PHYTOLACCACEAE
  Ball, Peter W.
Hardin, James W.
GYMNOSPERMAE (Carex) (Phytolacca)
Little, Elbert L. Jr. Carter, Richard
 
  (Cyperus) PIPERACEAE
CUPRESSACEAE Castaner, David
Bornstein, Allan J.
Adams, Robert P.
(Carex) Burger, William C.
(Juniperus) Core, Earl L.
St. John, Harold
Bartel, James A.
(Scleria)  
Vasek, Frank C.
Denton, Melinda F.
PITTOSPORACEAE
(Juniperus) (Cyperus) St. John, Harold
Zanoni, Thomas A.
Fairey, John E. III  
  (Scleria) PLANTAGINACEAE
EPHEDRACEAE Hermann, Frederick J.
Bassett, I. John
Benson, Lyman
(Carex) Rahn, Knud
  Jones, Stanley D.
 
PINACEAE Kral, Robert
PLUMBAGINACEAE
Critchfield, William B.
(Bulbostylis, Fimbristylis, Fuirena) Luteyn, James L.
Charlet, David Mastrogiuseppe, Joy
 
  (Carex) POACEAE
ANGIOSPERMAE Naczi, Robert F.C.
Aiken, Susan G.
  (Carex) (Festuca)
ACANTHACEAE Reznicek, Anton A.
Allred, Kelly W.
Thomas F. Daniel (Carex) (Aristida)
Meagher, Martha B.
Schuyler, Alfred E.
Aulbach-Smith, Cynthia A.
Wasshausen, Dieter C.
(Scirpus and segregate genera) (Chloris, Eustachys)
 

Smith, S. Galen

Barkworth, Mary E.
  (Eleocharis, Scirpus, and segregate genera)Baum, Bernard R.
ACERACEAE Strong, Mark
Beetle, Alan A.
Delendick, Thomas J.
Svenson, Henry K.
von Bothmer, Roland
  (Eleocharis) (Hordeum)
ACORACEAE Thieret, John W.
Brandenburg, David
Thompson, Sue (Cyperus) Brunken, Jere
  Tucker, Gordon C.
Campbell, Julian
AGAVACEAE (Cyperus) (Elymus)
Gentry, Howard S Zika, Peter
Carlson, Jack R.
Laurie , Robbins, R.
  (Pseudoroegneria)
(Yucca) CYRILLACEAE Cayouette, Jacques
Verhoek-Williams, Susan E.
Thomas, Joab L.
(Arctophila, Dupontia, Schizachne, Vahlodea)
    Church, George
ALISMATACEAE
DIAPENSIACEAE Crampton, Beecher
Beal, Ernest O.
Primack, Richard B.
Crins, William J.
(Sagittaria) Tiffney, Wesley N.
(Alopecurus, Milium, Ventenata)
Wooten, Jean W.
  Darbyshire, Stephen J.
  DROSERACEAE (Arctophila, Danthonia, Dupontia, Schizachne, Vahlodea)
AMARANTHACEAE
Donald E. Schnell Hal R. DeSelm
Mears, James A.
  (Andropogon, Bothriochloa, Dichanthelium, Schizachyrium)
  EBENACEAE de Wet, James
(Gomphrenoideae) White, Frank
(Bothriochloa, Cynodon, Dichanthium, Echinochloa,
Reed, Clyde F.
       Eleusine, Pennisetum, Setaria, Sorghum, Triticum,
Sauer, Jonathan D.
EMPETRACEAE      Zea, Zizania)
(Amaranthus) Eshbaugh, W. Hardy
Dewey, Douglas R.
    (Triticeae)
ANACARDIACEAE ERICACEAE Freckmann, Robert W.
Gillis, William T.
Dudley, Theodore R.
(Dichanthelium, Panicum)
Young, David A.
(Rhododendron) Greene, Craig W.
  Ebinger, John E.

(Calamagrostis)
ANNONACEAE (Kalmia) Guala, Gerald F. II
Kral, Robert
Hardin, James W.
(Neyraudia)
(Asimina, Deeringothamnus) (Kalmia) Harvey, LeRoy H.
  Hickman, James C.
Harvey, M.J.
APIACEAE (Menziesia) Holmgren, Arthur H.
Constance, Lincoln
Kloet, Sam P. Vander Kellogg, Elizabeth A.
Hartman, Ronald L.
(Vaccinium) (Poa)
  Krahn, Kathy
Koch, Stephen D.
APOCYNACEAE Prince, Martha
Krajina, Vladimir J.
Norman, Eliane M.
(Rhododendron) (Festuca, Vulpia)
(Vallesia) Skinner, H.T.
Laing, Charles C.
  (Rhododendron) (Ammophila)
AQUIFOLIACEAE Stevens, Peter F.
Lonard, Robert
Clark, Robert B.
Uttal, Leonard J.
(Vulpia)
Clark, Ross C.
(Vaccinium) Pavlick, Leon E.
Dudley, Theodore R.
Wells, Philip V.
(Festuca)
Krakow, Gregory A.
(Arctostaphylos) Peterson, Paul
(Ilex) Wilbur, Robert L.
Pyrah, Grant L.
    (×Agroelymus, ×Agrohordeum, Agropyron, ×Agrositanion)
ARACEAE ERIOCAULACEAE Reeder, Charlotte
Cornellier, Normand
Kral, Robert
Reeder, John R.
Huttleston, Donald G.
Moldenke, Harold N.
Riggins, Rhonda
Nicolson, Dan H.
  (Sporobolus)
Thompson, Sue A.
ERYTHROXYLACEAE Rogers, Blaine D.
  Plowman, Timothy C.
(Puccinellia)
ARALIACEAE   Scholz, Hildemar
St. John, Harold
EUPHORBIACEAE (Bromus)
  Burch, Derek
Snow, Neil
ARECACEAE Huft, Michael J.
(Diplachne, Hawaiian Grasses)
Glassman, Sidney F.
Ingram, John
Soreng, Robert J.
Hodel, Donald R.
Mayfield, Mark H.
(Poa)
Moore, Harold E. Jr. Webster, Grady L.
Spellenberg, Richard W.
Robert W. Read   Stieber, Michael T.
  FABACEAE (Ichnanthus)
ARISTOLOCHIACEAE Barneby, Rupert C.
Terrell, Edward E.
Pfeifer, Howard W.
(Astragalus, Cassia, Dalea, Parryella, Psorothamnus,  (Aegilops, Arrhenatherum, Dactylis, Hydrochloa, Lolium,
       Rhynchosia, Sophora, Sphinctospermum)      Luziola, Rhynchelytrum, Sorghum, Triticum)
ASCLEPIADACEAE Clewell, Andre F.
Wilson, F. Douglas
Drapalik, Donald J.
(Kummerowia, Lespedeza) (Sitanion)
Sundell, Eric
Duncan, Wilbur H.
Wipff, Joseph Karl
(Cynanchum) (Galactia) (Taeniatherum)
  Dunn, David
Witherspoon, John T.
ASTERACEAE (Lupinus) (Eragrostis)
Anderson, Dennis M.
Fantz, Paul R.
 
(Silphium) (Centrosema, Clitoria)
Anderson, Loran C.
Gillett, John M.
PODOSTEMACEAE
(Arnoglossum, Bigelowia, Chrysothamnus) (Trifolium) Wooten, Jean W.
Barbe, G. Douglas
Grant, William F.
 
(Cirsium) (Lotus)  
Barkley, Theodore M.
Gunn, Charles R.
POLEMONIACEAE
(Packera, Senecio) Hall, David W.
Day, Alva G.
Bayer, Randall J.
(Galactia) (Allophyllum, Collomia, Gilia, Gymnosteris, Ipomopsis,
(Antennaria) Isely, Duane
     Langloisia, Linanthus, Loeselia, Microseris, Navarretia)
Beetle, Alan A.
Nicolson, Dan H.
Timbrook, Steven L.
(Artemisia) (Desmodium) (Langloisia, Loeseliastrum)
Bierner, Mark W.
Riefner, Richard E. Jr. Wherry, Edgar T.
(Amblyolepis, Balduina, Dugaldia, Gaillardia, (Crotalaria) (Phlox)
     Helenium, Hymenoxys, Plummera) Rudd, Velva E.
Wilken, Dieter H.
Brown, Gregory K.
Ruff, William R.
 
(Platyschkuhria) (Strophostyles) POLYGALACEAE
Canne-Hilliker, Judith M.
Schubert, Bernice G.
Smith, Robert R.
(Galinsoga) (Desmodium) Wendt, Thomas L.
Carr, Gerald D.
Skinner, Steven G.
 
(Calycadenia, Dubautia, Osmadenia) (Crotalaria) POLYGONACEAE
Clark, W. Dennis
Small, Ernest
Hickman, James C.
(Hazardia) (Medicago) (Polygonum, Rumex)
Clonts, Jerry A.
Welsh, Stanley L.
Horton, James H.
(Elephantopus) (Hedysarum, Oxytropis) (Polygonella)
Coleman, James
Wiersema, John H.
Howard, Richard A.
(Verbesina) Windler, Donald R.
(Coccoloba)
Constance, Lincoln
(Crotalaria, Neptuna) Mitchell, Richard S.
(Eriophyllum) Wunderlin, Richard P.
(Polygonum, Rumex)
Cronquist, Arthur
(Bauhinia) Reveal, James L.
Davis, William S.
  (Aristocapsa, Centrostegia, Chorizanthe, Dedeckera,
(Lacamothrix, Malacothrix) FAGACEAE      Eriogonum, Gilmania, Hollisteria, Lastarriaea, Mucronea,
DeJong, D.C.D.
Duncan, Wilbur H.
     Mucronea, Nemacaulis, Oxytheca, Pterostegia,
(Astranthium) Guthrie, Roland L.
     Stenogonum, Systenotheca)
Douglas, George W.
(Quercus)  
(Antennaria, Arnica) Hardin, James W.
PONTEDERIACEAE
Ediger, Robert I.
Tucker, John M.
Horn, Charles N.
(Senecio) (Quercus) Lowden, Richard M.
Estes, James R.
  (Pontederia)
(Artemisia) FLACOURTIACEAE Wooten, Jean W.
Fisher, T. Richard
Keating, Richard
 
(Heliopsis, Silphium)   PORTULCACEAE
Gardner, Robert
FRANKENIACEAE Hohn, Janet
(Lipochaeta) Whalen, Molly A.
(Lewisia)
Gottlieb, Leslie D.
  Lewis, Walter H.
(Stephanomeria) FUMARIACEAE (Claytonia, Montia)
Harms, Vernon L.
Ownbey, Gerald B.
McNeill, John
(Heterotheca, Petasites) (Corydalis, Dicentra) Matthews, James F.
Hartman, Ronald L.
  (Portulaca)
(Machaeranthera) GARRYACEAE Ware, Stewart
Charles B. Heiser, Jr. Dahling, Gerald
(Talinum)
(Helianthus)    
Holmes, Walter C.
GENTIANACEAE POTAMOGETONACEAE
(Mikania) Broome, C. Rose
Haynes, Robert R.
Jackson, Raymond C.
(Centaurium) (Potamogeton)
(Iva) Gillett, John M.
 
Jansen, Robert K.
Maas, P.J.M.
PRIMULACEAE
(Spilanthes) (Voyria) Cooperrider, Tom S.
Johnson, Miles F.
Ornduff, Robert
Kelso, Sylvia
(Ageratum, Eupatorium) (Nymphoides)  
Johnson, R. Roy
Perry, James D.
PYROLACEAE
(Porophyllum) (Sabatia) Haber, Erich
Jones, Almut G.
Post, Douglas M.
 
(Aster, Brachyactis, Machaeranthera, Xylorhiza)  Pringle, James S.
RANUNCULACEAE
Jones, Samuel B. Jr. Yeo, Peter F.
Benson, Lyman
(Vernonieae)   (Ranunculus)
Keil, David J.
GESNERIACEAE Dennis, W. Michael
(Pectis) St. John, Harold (Clematis)
Koch, Rudy G.
(Hawaiian taxa) Duncan, Thomas
(Bidens) Skog, Laurence E.
(Ranunculus)
Kyhos, Donald W.
(Puerto Rican taxa) Keener, Carl S.
(Chaenactis)   Ramsey, Gwynn W.
Lamboy, Warren F.
GOODENIACEAE (Cimicifuga)
(Aster) St. John, Harold Scott, Peter J.
Lane, Meredith A.
  (Ranunculus)
(Gutierrezia, Gymnosperma, Xanthocephalum) GROSSULARIACEAE  
Lowrey, Timothy K.
Timbrook, Steven L.
RHAMNACEAE
(Tetramolopium) (Ribes) Johnston, Marshall C.
William F. Mahler   Schmidt, Clifford L.
(Baccharis) GUNNERACEAE (Ceanothus)
McGregor, Ronald L.
Palkovic, Lawrence A.
 
(Echinacea) St. John, Harold
ROSACEAE
Maguire, Bassett
  Davis, Hannibal A.
(Arnica) HALORAGACEAE (Rubus)
Mears, James A.
Aiken, Susan G.
Davis, Tyreeca S.
(Parthenium) (Myriophyllum) (Rubus)
Moe, L. Maynard
Catling, Paul M.
Johnston, Barry C.
(Matricaria) St. John, Harold
(Argentina, Comarum, Pentaphylloides, Potentilla,
Mooring, John
Wooten, Jean W.
     Sibbaldiopsis)
(Eriophyllum)   Ladd, Douglass
Morefield, James D.
HIPPOCASTANACEAE (Crataegus)
(Filagininae) Hardin, James W.
Lance, Ron
Nagata, Kenneth M.
  (Crataegus)
(Bidens) HYDRANGEACEAE Lewis, Walter H.
Neher, Robert T.
Hardin, James W.
(Rosa)
(Tagetes) (Hydrangea) Mellichamp, T. Lawrence
Nesom, Guy L.
  (Aruncus)
Nicolson, Dan H.
HYDROCHARITACEAE Morse, Larry E.
(Emilia) Cook, Christopher D.K.
(Rubus)
Niles, Wesley E.
St. John, Harold
Robertson, Kenneth R.
(Perityle)   Zika, Peter
Northington, David K.
HYDROPHYLLACEAE (Cotoneaster)
(Pyrrhopappus) Constance, Lincoln
 
Olsen, John S.
Howell, John T.
RUBIACEAE
(Verbesina) (Phacelia) Dempster, Lauramay T.
Ornduff, Robert
Lee, Gregory J.
(Galium)
(Lasthenia) (Phacelia) Kirkbride, Joseph H. Jr.
Ownbey, Gerald B.
  Lawson, Cheryl A.
(Cirsium) IRIDACEAE (Galium)
Parks, James C.
Goldblatt, Peter
Lewis, Walter H.
(Melanthera) Henderson, Douglass M.
(Diodia, Hedyotis, Richardia)
Parnell, Dennis R.
(Sisyrinchium) Nesom, Guy L.
(Corethrogyne) Hornberger, Kathleen L.
Terrell, Edward E.
Pinkava, Donald J.
(Sisyrinchium)  (Hedyotis, Houstonia, Oldenlandia, Oldenlandiopsis)
Pippen, Richard W.
Metcalf, Homer  
(Cacalia)   RUTACEAE
Powell, A. Michael
JUGLANDACEAE Bailey, Virginia L.
(Flaveria, Perityle) Manning, Wayne E.
(Ptelea)
Richards, Adrian J.
  Porter, Duncan M.
(Taraxacum) JUNCACEAE St. John, Harold
Richards, Edward L.
Ebinger, John E.
Scora, Rainer W.
(Ratibida) (Luzula) (Citrus, Fortunella)
Robinson, Harold E.
Hämet-Ahti, Leena
Stone, Benjamin C.
Schilling, Edward E.
Hermann, Frederick J.
 
Semple, John C.
(Juncus) SALICACEAE
(Aster, Borrichia, Chrysopsis, Heterotheca, Zika, Peter
Argus, George W.
     Pityopsis, Solidago, Xanthisma)   (Salix)
Shultz, Leila M. KRAMERIACEAE Dorn, Robert D.
(Artemisia) Simpson, Beryl B.
(Salix)
Simpson, Beryl B.
  Eckenwalder, James E.
(Acourtia) LAMIACEAE (Populus)
Smith, Edwin B.
Ball, Peter W.
 
Stebbins, G. Ledyard
Bozeman, John R.
SAPINDACEAE
(Antennaria, Crepis) (Dicerandra) St. John, Harold
Stoutamire, Warren P.
Cantino, Philip D.
 
(Gaillardia) (Physostegia) SAPOTACEAE
Strother, John L.
Collins, Joseph L.
Pennington, T.D.
Stuessy, Tod F.
(Scutellaria) St. John, Harold
Tanowitz, Barry D.
Irving, Robert S.
 
(Hemizonia) Lewis, F. Harlan
SARRACENIACEAE
Taylor, Constance E.S.
(Trichostema) Bell, C. Ritchie
(Euthamia, Solidago) Nelson, John B.
Case, Frederick W. II
Tomb, A. Spencer
Scora, Rainer W.
Schnell, Donald E.
(Lygodesmia, Stephanomeria) (Monarda)  
Torres, Andrew M.
Tucker, Arthur O.
SAXIFRAGACEAE
Tyrl, Ronald J.
(Coleus, Lavandula, Mentha, Origanum) Elvander, Patrick E.
(Achillea)   Mellichamp, T. Lawrence
Urbatsch, Lowell E.
LAURACEAE Ornduff, Robert
(Ericameria, Haplopappus, Tetragonotheca) Bergh, Robert
Packer, John G.
Van Horn, Gene S.
(Persea) (Chrysosplenium)
(Pentachaeta) van der Werff, Henk
Soltis, Douglas E.
Watson, Thomas J. Jr.   (Sullivantia)
(Xylorhiza) LEMNACEAE Wells, Elizabeth F.
Weedon, Ronald R.
Armstrong, Wayne P.
(Heuchera)
(Bidens) Beal, Ernest O.
 
Wells, James R.
  SCROPHULARIACEAE
(Polymnia) LENTIBULARIACEAE Canne-Hilliker, Judith M.
Wilken, Dieter H.
Godfrey, Robert K.
(Agalinis, Aureolaria)
(Hulsea) (Utricularia) Carr, Gerald D.
  Schnell, Donald E.
(Pedicularis)
BALSAMINACEAE Taylor, Peter
Cooperrider, Tom S.
Ornduff, Robert
  Elisens, Wayne J.
  LILIACEAE (Holmgrenanthe, Mabrya, Maurandya) 
BETULACEAE Ambrose, John D.
Ezell, Wayland L.
Clausen, Knud E.
Browne, Edward T. Jr. (Mimulus)
Furlow, John J.
(Aletris) Heckard, Lawrence R.
Mazzeo, Peter M.
Duncan, Wilbur H.
(Antirrhinum, Castilleja, Chaenorrhinum, Cordylanthus,
  (Yucca)      Orthocarpus, Parentucellia)
BIGNONIACEAE Flory, Walter S. Jr. Musselman, Lytton
Gentry, Alwyn
(Cooperia, Crinum, Furcraea, Galanthus, Habranthus,  Shaw, Richard J.
       Hippeastrum, Leucojum, Lycoris, Narcissus,  (Scrophularia)
BIXACEAE      Pancratium, Zephyranthes) Straw, Richard M.
Keating, Richard
Freeman, John D.
(Chionophila, Keckiella, Nothochelone, Penstemon)
  (Trillium) Thieret, John W.
BORAGINACEAE Hardin, James W.
Vickery, Robert K. Jr.
Frohlich, Michael W.
(Erythronium) (Mimulus)
(Heliotropium) Hickman, James C.
Yeo, Peter F.
Higgins, Larry C.
(Veratrum) (Euphrasia)
(Cryptantha) Jernstedt, Judith A.
 
Hommersand, Francia
(Chlorogalum) SMILACACEAE
(Amsinckia) Lenz, Lee W.
Duncan, Wilbur H.
Richardson, Alfred
(Triteleia) (Smilax)
(Tiquilia) Macfarlane, Roger M.
 
  (Fritillaria) SOLANACEAE
BRASSICACEAE Meerow, Alan W.
Averett, John E.
Al-Shehbaz, Ihsan
Mingrone, Louis V.
D'Arcy, William G.
Bartgis, Rodney L.
(Allium) Eshbaugh, W. Hardy
(Arabis) Preece, Sherman J.
Heiser, Charles B. Jr.
Kruckeberg, Arthur R.
(Zigadenus) (Capsicum, Solanum)
Mulligan, Gerald A.
Sherman, Harry L.
Schilling, Edward E.
Price, Robert A.
(Hastingsia, Schoenolirion) (Solanum)
(Draba, Erysimum) Skinner, Mark W.
Sullivan, Janet R.
Rodman, James E.
(Lilium) (Physalis)
Rollins, Reed C.
Utech, Frederick H.
 
Shaw, Elizabeth A.
Wilbur, Robert L.
SPARGANIACEAE
Stuckey, Ronald L.
(Uvularia) Beal, Ernest O.
(Armoracia, Nasturium, Rorippa)   Haynes, Robert R.
  LIMNANTHACEAE  
BROMELIACEAE Ornduff, Robert STERCULIACEAE
Smith, Lyman B.
  Cristobal, Carmen L.
  LINACEAE Goldberg, Aaron
BUDDLEJACEAE Rogers, Claude M.
Saunders, Jan
Norman, Eliane M.
   
St. John, Harold
LOASACEAE STYRACACEAE
  Prigge, Barry A.
Reveal, James L.
BURSERACEAE Thompson, Henry J.
(Halesia)
Porter, Duncan M.
   
  LOGANIACEAE SYMPLOCACEAE
CACTACEAE St. John, Harold
Hardin, James W.
Anderson, Edward F.
   
Benson, Lyman
LYTHRACEAE TAMARICACEAE
R. Hunt, David Graham, Shirley A.
Baum, Bernard R.
Taylor, Nigel
  (Tamarix)
  MAGNOLIACEAE  
CALLITRICHACEAE James W. Hardin THEACEAE
Beal, Ernest O.
  Spongberg, Stephen A.
  MALPIGHIACEAE  
CALYCANTHACEAE Bruce MacBryde THYMELAEACEAE
Hardin, James W.
(Galphimia) Nevling, Lorin I. Jr. 
  Vivaldi, Jose L.
(Daphne, Daphnopsis, Dirca, Edgeworthia, Passerina)
CAMPANULACEAE   St. John, Harold
Hohn, Janet
MALVACEAE (Wikstroemia)
Shetler, Stanwyn G.
Fryxell, Paul A.
 
(Campanula) Hill, Steven R.
TURNERACEAE
    Arbo, Maria Mercedes
CANNABACEAE MELASTOMATACEAE Ornduff, Robert
Small, Ernest
Kral, Robert
 
  (Rhexia) TYPHACEAE
CAPPARACEAE Morley, Thomas
Beal, Ernest O.
Iltis, Hugh H.
(Mouriri) Smith, S. Galen
  Wurdack, John J.
 
CAPRIFOLIACEAE   URTICACEAE
Dudley, Theodore R. MELIACEAE Bassett, I. John
James, W. Hardin,  Pennington, T.D.
Boufford, David E.
Salamun, Peter J.
  Woodland, Dennis W.
  MONOTROPACEAE  
CARYOPHYLLACEAE Wallace, Gary D.
VALERIANACEAE
Crow, Garrett E.
  Meyer, Frederick G.
(Sagina) MYRTACEAE (Valeriana)
Hartman, Ronald L.
McVaugh, Rogers
Ware, Donna M.E.
Kruckeberg, Arthur R.
St. John, Harold
(Valerianella)
(Lychnis, Silene)    
McNeill, John
NAJADACEAE VERBENACEAE
Rabeler, Richard K.
Haynes, Robert R.
Moldenke, Harold N.
    Sanders, Roger W.
CELASTRACEAE NYCTAGINACEAE (Duranta, Lantana)
Prigge, Barry A.
Galloway, Leo A.
Umber, Ray E.
  Reed, Clyde F.
(Glandularia)
CERATOPHYLLACEAE Spellenberg, Richard W.
 
Les, Donald H.
  VIOLACEAE
Lowden, Richard M.
NYMPHAEACEAE McKinney, Landon E.
(Ceratophyllum) Beal, Ernest O.
Russell, Norman H.
Wooten, Jean W.
Wiersema, John H.
 
    VISCACEAE
CHENOPODIACEAE NYSSACEAE Kuijt, Job
Arnold, H. Lamont
Eyde, Richard H.
Wiens, Delbet
(Salsola)    
Ball, Peter W.
OLEACEAE VITACEAE
(Arthrocnemum, Salicornia, Sarcocornia) Little, Elbert L. Jr. Duncan, Wilbur H.
Barbe, G. Douglas
(Fraxinus) Moore, Michael O.
(Salsola)    
Bassett, I. John
ONAGRACEAE XYRIDACEAE
(Atriplex) Boufford, David E.
Kral, Robert
Blackwell, Will H. Jr. (Circaea)  
McNeill, John
Lewis, F. Harlan
ZANNICHELLIACEAE
(Atriplex) (Clarkia, Gayophytum) Beal, Ernest O.
Reed, Clyde F.
Straley, Gerald B.
 
Sanderson, Stewart C.
(Oenothera) ZINGIBERACEAE
Stutz, Howard
Towner, Howard
Smith, Rosemary M.
(Atriplex) (Calylophus)  
Wilson, Hugh D.
Wagner, Warren L.
ZYGOPHYLLACEAE
(Chenopodium) (Camissonia, Gaura, Oenothera) Porter, Duncan M.
     
CHRYSOBALANACEAE ORCHIDACEAE  
Prance, Ghillean T.
Brown, Paul Martin
 
  (North American native Orchids)  
CISTACEAE Catling, Paul M.
 
Morse, Larry E.
(Spiranthes)  
(Hudsonia) Dressler, Robert L.
 
Wilbur, Robert L.
(Encyclia, Epidendrum)  
  Luer, Carlyle A.
 
CLUSIACEAE Magrath, Lawrence K.
 
Adams, Preston
Meades, Susan
 
Stevens, Peter F.
Merhoff, Loyal A. III  
(Calophyllum) (Calypso, Platanthera)  
  Sheviak, Charles
 
COMMELINACEAE Snyder, David
 
Dunn, David
   
Faden, Robert B.
OROBANCHACEAE  
Hunt, David R.
Collins, Turner
 
MacRoberts, D.T.
(Orobanche)  

 

Repositories visited by BONAP or from which plant materials and data have been exchanged

Academy of Natural Sciences (Philadelphia, PA)
Acadia National Park (Bar Harbor, ME)
Acadia University (Wolfville, Nova Scotia, Canada)
Accokeek Foundation Herbarium, Piscataway Park (Accokeek, MD)
Ada Hayden Herbarium, Iowa State Universtiy (Ames, IA)
Agriculture and Agri-Food Canada (Charlottetown, Prince Edward Island, Canada)
Agriculture and Agri-Food Canada (Saskatoon, Saskatchewan, Canada)
Agriculture and Agri-Food Canada (Swift Current, Saskatchewan, Canada)
Alaska Region, USDA Forest Service (Sitka, AK)
Albertson College (Caldwell, ID)
Albion R. Hodgdon Herbarium, University of New Hampshire (Durham, NH)
Alison Cusick collections housed at University of North Carolina (Chapel Hill, NC)
Amherst College (Amherst, MA)
Andrew Charles Moore Herbarium, University of South Carolina (Columbia, SC)
Angelo State University (San Angelo, TX)
Anne Arundel Community College (Arnold, MD)
Anniston Museum of Natural History (Anniston, AL)
Arches National Park (Moab, UT)
Arizona State University (Tempe, AZ)
Arkansas Natural Heritage Commission
Arkansas State University (Jonesboro, AR)
Arkansas Tech University (Russellville, AR)
Arnold Arboretum, Harvard University (Cambridge, MA)
Asbury College (Wilmore, KY)
Austin Peay State University (Clarksville, TN)
Australian National Botanic Gardens (Canberra, Australia)
Bard College (Annandale-on-Hudson, NY)
Baylor University (Waco, TX)
Berea College (Berea, KY)
Bighorn National Forest (Sheridan, WY)
Biltmore Herbarium, housed at North Carolina State University (Raleigh, NC)
Birmingham Botanical Garden (Birmingham, AL)
Bodega Marine Laboratory, University of California Davis Affiliation (Davis, CA)
Botanical Garden and Museum (Oslo, Norway)
Botanical Museum (Lund, Sweden)
Botanical Research Institute of Texas (Fort Worth, TX)
Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitat Berlin (Berlin, Germany)
Botanisches Institut und Botanischer der Universität Bonn (Bonn, Germany)
Brigham Young University (Provo, UT)
British Columbia Conservation Data Centre computer databases (Victoria, British Columbia, Canada)
British Museum, The Natural History Museum (London, U.K.)
Brooklyn Botanic Garden (Brooklyn, NY)
Brown University (Providence, RI)
Bryce Canyon National Park (Bryce, UT)
Bucknell University (Lewisburg, PA)
Butler University (Indianapolis, IN)
C.E. Bessey Herbarium, University of Nebraska State Museum (Lincoln, NE)
C.L. Lundell Herbarium, University of Texas (Austin, TX)
California Academy of Sciences (San Francisco, CA)
California Department of Food and Agriculture (Sacramento, CA)
California State University (Chico, CA)
California State University (Fresno, CA)
California State University (Northridge, CA)
Campbellsville College (Campbellsville, KY)
Canadian Forest Service (Ottawa, Ontario, Canada)
Canadian Museum of Nature (Ottawa, Ontario, Canada)
Canisius College (Buffalo, NY)
Canyonlands National Park (Moab, UT)
Cape Cod Museum Natural-History
Capitol Reef National Park (Torrey, UT)
Carl Sharsmith Herbarium, San Jose State University (San Jose, CA)
Carnegie Museum of Natural History (Pittsburgh, PA)
Catholic University of America (Washington, DC)
Cedar Breaks National Monument (Cedar Breaks, UT)
Central Missouri State University (Warrensburg, MO)
Central Washington University (Ellensburg, WA)
Central Wyoming College (Riverton, WY)
Centre College of Kentucky (Danville, KY)
Chadron State College (Chadron, NE)
Chicago Botanic Garden (Gelncoe, IL)
Chrysler Herbarium, Rutgers University (New Brunswick, NJ)
Clark University (Worcester, MA)
Claude E. Phillips Herbarium, Delaware State College (Dover, DE)
Clemson University (Clemson, SC)
Cleveland Museum of Natural History (Cleveland, OH)
Clinton Herbarium, Buffalo Museum of Science (Buffalo, NY)
Coe College (Cedar Rapids, IA)
College of Environmental Sciences and Forestry, State University of New York (Syracuse, NY)
College of the Agriculture, University of Kentucky (Lexington, KY)
College of the Ozarks (Point Lookout, MO)
Colorado College (Colorado Springs, CO)
Colorado State University (Fort Collins, CO)
Connecticut Botanical Society, Yale University (New Haven, CT)
Connecticut College (New London, CT)
Cornell University (Ithaca, NY)
Cove Point Natural Heritage Trust (Lusby, MD)
Cove Point Natural Heritage Trust Herbarium (Prince Frederick, MD)
Cranberry Lake Biological Station (Syracuse, NY)
Crater Lake National Park Herbarium (Crater Lake, OR)
Creso Herbarium, Pacific Lutheran University (Tacoma, WA)
Davies Herbarium, University of Louisville (Louisville, KY)
Davis and Elkins College (Elkins, WV)
Deaver Herbarium, Northern Arizona University (Flagstaff, AZ)
Delaware Natural Heritage Program (Smyrna, DE)
Delta Institute of Natural History (Bowdoin, ME)
Delta State University (Cleveland, MS)
Denali National Park (Denali, AK)
Diatom Herbarium, Academy of Natural Sciences (Philadelphia, PA)
Dinosaur National Monument (Dinosaur, CO)
Disney Wilderness Preserve/The Nature Conservancy (Kissimmee, FL)
Douglas County Museum (Roseburg, OR)
Drury College (Springfield, MO)
Dudley Herbarium, California Academy of Sciences (San Francisco, CA)
Duke University (Durham, NC)
Dunn-Palmer Herbarium, University of Missouri (Columbia, MO)
E.L. Reed Herbarium, Texas Tech University (Lubbock, TX)
East Herbarium, National Capital Parks (Washington, DC)
Eastern Illinois University (Charleston, IL)
Eastern Kentucky University (Richmond, KY)
Eastern New Mexico University (Portales, NM)
Eastern Oregon University (La Grande, OR)
Economic Herbarium of Oakes Ames, Harvard University (Cambridge, MA)
Eglin Air Force Base (Okaloosa County, FL)
Emory University (Atlanta, GA)
Emporia State University (Emporia, KS)
Evergreen State College (Olympia, WA)
Fairbanks Museum and Planetarium (Saint Johnsbury, VT)
Fairchild Tropical Garden (Miami, FL)
Fan Memorial Institute of Biology (Beijing, China)
Fessenden Herbarium, Cylburn Arboretum (Baltimore, MD)
Field Museum of Natural History (Chicago, IL)
Florida Atlantic University (Boca Raton, FL)
Florida State University (Tallahassee, FL)
Forest Pathology Herbarium, Rocky Mountain Research Station, USDA Forst Service (Fort Collins, CO)
Forest Service Herbarium, Juneau Forestry Sciences Laboratory (Juneau, AK)
Forest Service Herbarium, Rocky Mountain Forest and Range Experiment Station (Fort Collins, CO)
Forest Service Region4, USDA Forest Service (Ogden, UT)
Forest Service, Southern Forest Experiment Station (Pineville, LA)
Fort Polk Herbarium (Fort Polk, LA)
Fort Worth Museum of Science and History (Fort Worth, TX)
Fossil Butte National Monument (Kemmerer, WY)
Friesner Herbarium, Butler University (Indianapolis, IN)
Frostburg State University (Frostburg, MD)
Furman University (Greenville, SC)
Garrett Herbarium, University of Utah (Salt Lake City, UT)
George Mason University (Fairfax, VA)
George R. Clooney Herbarium, Colgate University (Hamilton, NY)
Georgetown College (Georgetown, KY)
Grand Teton National Park (Moose, WY)
Gray Herbarium, Harvard University (Cambridge, MA)
Great Lakes Forestry Centre, Canadian Forest Service (Sault Sainte Marie, Ontario, Canada)
H.J. Andrews Experimental Forest (Corvallis, OR)
Harold L. Lyon Arboretum (Honolulu, HI)
Hastings Natural History Reservation (Carmel Valley, CA)
Henderson State University (Arkadelphia, AR)
Hendrix College (Conway, AR)
Herbarium Pacificum, Bishop Museum (Honolulu, HI)
Herbier du Québec, Complexe scientifique (Sainte-Foy, Quebec, Cananda)
Herbier Rolland-Germain, Universite de Sherbrooke (Sherbrooke, Quebec, Canada)
Houghton College (Houghton, NY)
Howard Payne University (Brownwood, TX)
Hoysradt Herbarium, Hartwick College (Oneonta, NY)
Humboldt State University (Arcata, CA)
Idaho State University (Pocatello, ID)
Illinois Natural History Survey (Champaign, IL)
Illinois State Museum (Springfield, IL)
Illinois State University (Normal, IL)
Indiana State University (Terre Haute, IN)
Indiana University (Bloomington, IN)
Indiana University Southeast (New Albany, IN)
Institut des Universitaires et Musée Lecoq (Clermont-Ferrand, France)
Institute for Botanical Exploration (Mississippi State, MS)
Intermountain Forest and Range Experiment Station, US Forest Service (Boise, ID)
Iowa Department of Natural Resources (Des Moines, IA)
Iowa Native Plant Society (IA)
Jacksonville State University (Jacksonville, AL)
Jacksonville State University (Jacksonville, AL)/Anniston Museum of Natural History (Anniston, AL)
Jamie Whitten Delta States Research Center, USDA/ARS (Stoneville, MS)
Jepson Herbarium, University of California (Berkeley, CA)
Jesup Herbarium, Dartmouth College (Hanover, NH)
John D. Freeman Herbarium, Auburn University (Auburn, AL)
Kagoshima University (Kagoshima, Japan)
Kansas State University (Manhattan, KS)
Kathryn Kalmbach Herbarium, Denver Botanic Gardens (Denver, CO)
Keene State College (Keene, NH)
Kent State University (Kent, OH)
Kentucky State Nature Preserves Commission collections housed at Eastern Kentucky University (Richmond, KY) or Davies Herbarium, University of Louisville (Louisville, KY)
Kew Gardens, Royal Botanic Society (London, U.K.)
Lambert C. Erickson Weed Diagnostic Laboratory, University of Idaho (Moscow, ID)
Laurentian University (Sudbury, Ontario, Canada)
Laval University (Sainte-Foy, Quebec, Canada)
Lindsey Wilson College (Columbia, KY)
Linfield College (McMinnville, OR)
Longwood University (Farmville, VA)
Lord Fairfax Community College (Middletown, VA)
Louisiana State University (Baton Rouge, LA)
Louisiana State University (Shreveport, LA)
Louisiana Tech University (Ruston, LA)
Lyndon State College (Lydonville, VT)
Mammoth Cave National Park (Mammoth Cave, KY)
Marie Selby Botanical Gardens (Sarasota, FL)
Marine Biological Laboratory, George M. Gray Museum (Woods Hole, MA)
Marion Ownbey Herbarium, Washington State University (Pullman, WA)
Marshall University (Huntington, WV)
Mary Wharton collections housed at University of Michigan (Ann Arbor, MI)
Massachusetts College of Liberal Arts (North Adams, MA)
Max Medley collections housed at Western Kentucky University (Bowling Green, KY)
McGill University (Sainte-Anne-de-Bellevue, Quebec, Canada)
McNeese State University (Lake Charles, LA)
Mecklenburg County Park and Recreation (Charlotte, KY)
Michigan State University (East Lansing, MI)
Middle Tennessee State University (Murfreesboro, TN)
Middlebury College (Middlebury, VT)
Milwaukee Public Museum (Milwaukee, WI)
Minnesota Department of Natural Resources (St. Paul, MN)
Mississippi Museum of Natural Science (Jackson, MS)
Mississippi State University (Mississippi State, MS)
Missouri Botanical Garden (St. Louis, MO)
Mohr Herbarium, Alabama Museum of Natural History (Tuscaloosa, AL)
Mohonk Preserve (New Paltz, NY)
Monroe County Department of Parks (Rochester, NY)
Montana State University (Bozeman, MT)
Montreal Botanical Garden Herbarium (Montreal, Quebec, Canada)
Moorhead State University (Moorhead, MN)
Morehead State University (Morehead, KY)
Morris Arboretum, University of Pennsylvania (Philadelphia, PA)
Morton Arboretum (Lisle, IL)
Mount Rainier National Park (Ashford, WA)
Murray State University (Murray, KY)
Museum National d'Histoire Naturelle (Paris, France)
Muskingum College (New Concord, OH)
N. F. Flynn Memorial Herbarium, Goddard College (Plainfield, VT)
Nantucket Maria Mitchell Association (Nantucket, MA)
National Tropical Botanical Garden (Lawai, HI)
Natural Bridges National Monument (Lake Powell, UT)
Natural History Museum of Los Angeles County (housed at RSA, Claremont, CA)
Natural History Museum of Los Angeles County (Los Angeles, CA)
Nebraska Natural Heritage Herbarium (Lincoln, NE)
Nevada State Museum (Carson City, NV)
New Brunswick Museum (Saint John, New Brunswick, Canada)
New England Botanical Club, Harvard University (Cambridge, MA)
New England College (Henniker, NH)
New England Wild Flower Society (Framingham, MA)
New Mexico State University (Las Cruces, NM)
New York Botanical Garden (Bronx, NY)
New York State Museum (Albany, NY)
Nicholls State University (Thibodaux, LA)
North Carolina State University (Raleigh, NC)
North Coast Herbarium of California, Sonoma State University (Rohnert Park, CA)
North Dakota State University (Fargo, ND)
Northeast Louisiana University (Monroe, LA)
Northern Illinois University (DeKalb, IL)
Northern Kentucky University (Highland Heights, KY)
Northern Michigan University (Marquette, MI)
Norton-Brown Herbarium, University of Maryland (College Park, MD)
Nova Scotia Museum of Natural History (Halifax, Nova Scotia, Canada)
Ohio State University (Columbus, OH)
Oklahoma State University (Stillwater, OK)
Old Dominion University (Norfolk, VA)
Olga Lakela Herbarium, University of Minnesota Duluth (Duluth, MN)
Olympic National Park (Port Angeles, WA)
Oregon Flora Project (Corvallis, OR)
Oregon Natural Heritage Information Center (Corvallis, OR)
Oregon State University (Corvallis, OR)
Ozarks Regional Herbarium, Southwest Missouri State University (Springfield, MO)
Pacific Gas and Electric Company Herbarium (San Ramon, CA)
Pacific Grove Museum of Natural History (Pacific Grove, CA)
Pacific Union College (Angwin, CA)
Patuxent Wildlife Research Center Herbarium (Laurel, MD)
Peabody Essex Museum (Salem, MA)
Pennsylvania State University (University Park, PA)
Philadelphia Scout Ranch Herbarium (PA)
Pine Mountain Settlement School (Pine Mountain, KY)
Planting Fields Arboretum (Oyster Bay, NY)
Plummer's Island Collection, housed at Smithsonian Institution (Washington, DC)
Point Lobos State Reserve Herbarium
Pomona College (housed at RSA, Claremont, CA)
Prince Rupert Forest Region (Smithers, British Columbia, Canada)
Pringle Herbarium, University of Vermont (Burlington, VT)
Private herbarium collection of B.A. Bennett (Whitehorse, YT)
Private herbarium collection of C. Davis
Private herbarium collection of C. Fleming
Private herbarium collection of Edna Banta (Bloomington, IN)
Private herbarium collection of Eric Hulten (Lund, Sweden)
Private herbarium collection of F. Hirst
Private herbarium collection of Frank C. Seymour
Private herbarium collection of G.F. Ledingham (Regina, Saskatchewan, Canada)
Private herbarium collection of Jim Vanderhorst (Elkins, WV)
Private herbarium collection of John Schwegman
Private herbarium collection of Johnnie Varner (Cynthiana, KY)
Private herbarium collection of M. Oldham (Ontario, Canada)
Private herbarium collection of Pamela Weatherbee
Private herbarium collection of Paul J. Harmon (Elkins, WV)
Private herbarium collection of R. Bartgis
Private herbarium collection of R. Wilson
Private herbarium collection of R.H. Waldo
Private herbarium collection of Ralph M. Kriebel (Bedford, IN)
Private herbarium collection of Robert H. Mohlenbrock (Carbondale, IL)
Private herbarium collection of Rodney L. Bartgis
Private herbarium collection of Scott Shriver, Albert Shriver, and Clete Smith (Pittsburg, PA)
Private herbarium collection of Thomas Rawinski
Private herbarium collection of W.D. McIlveen
Private herbarium collection of William Grafton
Pullen Herbarieum, University of Mississippi (University, MS)
Purdue University (West Lafayette, IN)
Purdue University Indianapolis (Indianapolis, IN)
Queens College (Queens, NY)
Queen's University (Kingston, Ontario, Canada)
R.L. McGregor Herbarium, University of Kansas (Lawrence, KS)
R.M. Myers Herbarium, Western Illinois University (Macomb, IL)
Ramsey-Freer Herbarium, Lynchburg College (Lynchburg, VA)
Rancho Santa Ana Botanic Garden (Claremont, CA)
Range Science Herbarium, New Mexico State University (Las Cruces, NM)
Raymond Athey Herbarium, University of Memphis (Memphis, TN)
Reed College (Portland, OR)
Reed Herbarium (housed at MO, St. Louis, MO)
Rice Creek Field Station, State University of New York-Oswego (Oswego, NY)
Richmond Nature Park (Richmond, British Columbia, Canada)
Robert A. Vines Environmental Science Center (Houston, TX)
Robert F. Hoover Herbarium, California Polytechnic State Univeristy (San Luis Obispo, CA)
Robert W. Freckmann Herbarium, University of Wisconsin at Stevens Point (Stevens Point, WI)
Rochester Academy of Science (Rochester, NY)
Rocky Mountain Forest and Range Experiment Station (Albuquerque, NM)
Rocky Mountain Herbarium, University of Wyoming (Laramie, WY)
Rocky Mountain Herbarium, University of Wyoming (Laramie, WY) housed at Forest Service Herbarium, Rocky Mountain Forest and Range Experiment Station (Fort Collins, CO)
Rocky Mountain Research Station (Provo, UT)
Ross Clark collection from Red River Gorge housed at Eastern Kentucky University (Richmond, KY)
Royal Botanical Gardens (Hamilton, ON)
Royal British Columbia Museum (Victoria, British Columbia, Canada)
S.M. Tracy Herbarium, Texas A&M University (College Station, TX)
Sagehen Creek Field Station Herbarium (Truckee, CA)
Saint Bonaventure University (Saint Bonaventure, NY)
Saint Lawrence University (Canton, NY)
Salem College (Winston-Salem, NC)
Salem International University (Salem, WV)
Sam Houston State University (Huntsville, TX)
San Diego Natural History Museum (San Diego, CA)
San Francisco State University (San Francisco, CA)
San Juan College (Farmington, NM)
Santa Barbara Museum of Natural History (Santa Barbara, CA)
Santa Rosa Junior College (Santa Rosa, CA)
Scarborough College, University of Toronto (Toronto, Ontario, Canada)
Seymour Herbarium (Burlington, VT)
Shawnee National Forest (Harrisburg, IL)
Slater Museum of Natural History, University of Puget Sound (Tacoma, WA)
Snake River Plains Herbarium, Boise State University (Boise, ID)
South Dakota State University (Brookings, SD)
South Florida Collections Management Center, Everglades National Park (Homestead, FL)
Southeast Missouri State University (Cape Girardeau, MO)
Southern Illinois University (Carbondale, IL)
Southern Methodist University (Dallas, TX)
Southern Oregon University (Ashland, OR)
St. John's College (Cambridge, England)
St. John's University/College of Saint Benedict (Collegeville, MN)
St. Mary's College (Notre Dame, IN)
State University of New York, College at Oneonta (Oneonta, NY)
Staten Island Institute of Arts and Science (Staten Island, NY)
Stephen F. Austin State University (Nacogdoches, TX)
Sul Ross State University (Alpine, TX)
Swedish Museum of Natural History (Stockholm, Sweden)
Sweeny Granite Mountains Deser Research Center
Syracuse University (Syracuse, NY)
Tawes State Office Building (Annapolis, MD)
Texas A&M University (College Station, TX)
The College of William and Mary (Williamsburg, VA)
Togiak National Wildlife Refuge (Dillingham, AK)
Towson State University (Baltimore, MD)
Troy University (Troy, AL)
Truman G. Yuncker Herbarium, DePauw University (Greencastle, IN)
Tulane University (New Orleans, LA)
U.S. Bureau of Land Management (Lakeview, OR)
U.S. Bureau of Land Management (Painted Rock Ranch, CA)
U.S. Geological Survey, Lafayette Wetlands Center (Lafayette, LA)
UC Davis Herbarium, University of California (Davis, CA)
United States National Arboretum, USDA/ARS (Washington, DC)
United States National Herbarium, Smithsonian Institution (Washington, DC)
Universidad Nacional Autónoma de México (Mexico City, Mexico)
Universite Laval (Sainte-Foy, Quebec, Canada)
University of Alabama (Tuscaloosa, AL)
University of Alaska (Fairbanks, AK)
University of Alberta (Edmonton, Alberta, Canada)
University of Arizona (Tucson, AZ)
University of Arkansas (Fayetteville, AR)
University of Arkansas (Monticello, AR)
University of Arkansas at Little Rock (Little Rock, AR)
University of British Columbia (Vancouver, British Columbia, Canada)
University of Calgary (Calgary, Alberta, Canada)
University of California (Berkeley, CA)
University of California (Davis, CA)
University of California (Irvine, CA)
University of California (Los Angeles, CA)
University of California (Santa Barbara, CA)
University of California (Santa Cruz, CA)
University of California at Riverside (Riverside, CA)
University of Charleston (Charleston, WV)
University of Cincinnati (Cincinnati, OH)
University of Colorado (Boulder, CO)
University of Connecticut (Storrs, CT)
University of Florida Herbarium, Florida Museum of Natural History (Gainesville, FL)
University of Georgia (Athens, GA)
University of Guelph (Guelph, Ontario, Canada)
University of Hawaii (Honolulu, HI)
University of Idaho (Moscow, ID)
University of Idaho, College of Forestry (Moscow, ID)
University of Illinois (Urbana, IL)
University of Iowa (housed at ISC, Ames, IA)
University of Kentucky (Lexington, KY)
University of Maine (Orono, ME)
University of Maryland (College Park, MD)
University of Massachusetts (Amherst, MA)
University of Michigan (Ann Arbor, MI)
University of Minnesota (St. Paul, MN)
University of Minnesota Herbarium, J.F. Bell Museum of Natural History (St. Paul, MN)
University of Montana (Missoula, MT)
University of Montreal (Montreal, Quebec, Canada)
University of Nebraska (Omaha, NE)
University of Nebraska at Kearney (Kearney, NE)
University of Nevada (Las Vegas, NV)
University of Nevada (Reno, NV)
University of New Brunswick (Fredericton, New Brunswick, Canada)
University of New Mexico (Albuquerque, NM)
University of New Orleans (New Orleans, LA)
University of North Alabama (Florence, AL)
University of North Carolina (Chapel Hill, NC)
University of Northern Colorado (Greeley, CO)
University of Notre Dame (Notre Dame, IN)
University of Oklahoma (Norman, OK)
University of Oregon (housed at OSC, Corvallis, OR)
University of Patras (Patras, Greece)
University of Pennsylvania (Philadelphia, PA)
University of Peshawar (Peshawar, Pakistan)
University of Prince Edward Island (Charlottetown, Prince Edward Island, Canada)
University of Rhode Island (Kingston, RI)
University of the South (Sewanee, TN)
University of South Alabama (Mobile, AL)
University of South Florida (Tampa, FL)
University of Southern Mississippi (Hattiesburg, MS)
University of Southwestern Louisiana (Lafayette, LA)
University of Tennessee (Knoxville, TN)
University of Texas (Austin, TX)
University of Texas (El Paso, TX)
University of Victoria (Victoria, British Columbia, Canada)
University of Washington (Seattle, WA)
University of Waterloo (Waterloo, Ontario, Canada)
University of West Florida (Pensacola, FL)
University of Western Ontario (London, Ontario, Canada)
University of Wisconsin (La Crosse, WI)
University of Wisconsin (Madison, WI)
University of Wisconsin (Superior, WI)
University of Wisconsin at Green Bay (Green Bay, WI)
University of Wisconsin at Oshkosh (Oshkosh, WI)
University of Wisconsin at Platteville (Platteville, WI)
University of Wyoming (Laramie, WY)
Universtiy of Northern Iowa (Cedar Falls, IA)
US Forest Service Herbarium (NV)
Utah State University (Logan, UT)
Utah Valley University (Orem, UT)
V. L. Komarov Botanical Institute, Russian Academy of Sciences (Saint Petersburg, Russia)
Valdosta State University (Valdosta, GA)
Vanderbilt University (Nashville, TN)
Vascular Plant Herbarium, Agriculture and Agri-Food Canada (Ottawa, Ontario, Canada)
Vascular Plant Herbarium, Royal Ontario Museum (Toronto, Ontario, Canada)
Virginia Natural Heritage Program (Richmond, VA)
Virginia Polytechnic Institute and State University (Blacksburg, VA)
W.P. Fraser Herbarium, University of Saskatchewan (Saskatoon, Saskatchewan, Canada)
W.S. Turrell Herbarium, Miami University (Oxford, OH)
Wabash College (Crawfordsville, IN)
Walter B. McDougall Herbarium, Museum of Northern Arizona (Flagstaff, AZ)
Washington State University (Pullman, WA)
Weeks Bay Herbarium
West Texas A&M University (Canyon, TX)
West Virginia Army National Guard at Camp Dawson (Camp Dawson, WV)
West Virginia Natural Heritage Program (Elkins, WV)
West Virginia University (Morgantown, WV)
Western Carolina University (Cullowhee, NC)
Western Kentucky University (Bowling Green, KY)
Western Washington University (Bellingham, WA)
Westfield State College (Westfield, MA)
Weston New Mexico University (Silver City, NM)
Whitman College (Walla Walla, WA)
Wilfrid Laurier University (Waterloo, Ontario, Canada)
Willamette University, housed at Oregon State University (Corvallis, OR)
William Darlington Herbarium, West Chester University (West Chester, PA)
William Penn University (Oskaloosa, IA)
Windham College (Putney, VT)
Wrangell-St. Elias National Park and Preserve (Copper Center, AK)
Yale University (New Haven, CT)
Yellowstone National Park (Yellowstone National Park, WY)
Youngstown State University (Youngstown, OH)


Field botanists

Abel, William Eastman, Don Jones, Almut G. Neuman, Ray Snyder, David
Aiken, Susan G. Eastman, Priscilla A. Jones, Stanley D. Nordman, Carl Soreng, Robert J.
Alford, Mac Eckel, P.M. Juelg, Tussell Norquist, Cary Sorrie, Bruce
Allen, Charles Ehrenfeld, Joan G. Kauffman, Randy Norton,, Dan H. Spaulding, Dan
Allison, James Ellis, Donna Kaul, Robert B. Nunn, Sarah Spellenberg, Richard
Allred, Kelly W. Emens, Rob Kearns, Kelly O'Brien, Kate Sperduto, Dan
Alverson, Ed Erickson, Terrell Keener, Carl O'Kane, Steve L. Sprinkle, Danny
Anderson, Loran C. Ernst, John A. Keith, Eric O'Kennon, Robert Starr, Forest
Angelo, Ray Estes, Dwayne Ketchersid, Mary Ode, David J. Stensvold, Mary Clay
Argus, George W. Evans, Jeffrey Kinter, Cecilia L. Old, Patrick Steury, Brent
Aulbach-Smith, Cynthia Fenner, Patti Kral, Robert Old, Richard Stewart, Leslie
Ault, James R. Fertig, Walter Kratz, Andrew Oldham, Michael Stohlgren, Tom
Austin, Daniel F. Fisler, Deborah Kunzer, John M. Osorio, Federico G. Stratford, Kay
Aylor, Bill Ford-Werntz, Donna Kyde, Kerrie L. Otsuki, Yuka Sundberg, Scott
Barger, Wayne Forestieri, David Lance, Ron Packer, Pat Swearingen, Jil
Barker, William Fortney, Ron Langdon, K. Palumbo, Theresa Talbot, Steve
Barkley, Theodore M. Fowler, Larry Larson, Gary Parsons, Jenifer Terrell, Edward E.
Barkworth, Mary E. Franklin, Ben Lass, Larry Peet, Robert Tessene, Paul
Barnes, Bruce Frantz, Paul Lata, Mary Penskar, Michael Thieret, John W.
Bartgis, Rodney L. Freeman, Craig C. Lawson, Cheryl A. Perkins, Kent D. Thomas, L.K.
Baum, Bernard French, John R. Lederer, Nancy Peterson, Angela Thompson, Sue
Bayer, Randall J. Frey, Mark Leonard, Richard Peterson, Paul Thurmond, M.
Bennett, Bruce Frye, Chris Leonard, Steven W. Piep, Michael Till, Linda
Bishop, Andrea Fryxell, Paul A. Lesica, Peter Pinzl, Ann Tolley, Regan
Bissell, Jim Furqueron, Chris Lichvar, Bob Pittillo, Dan Towsend, Lynn
Bjork, McIntosh Gage, Sarah Lipkin, Rob Pollack, Glenn Trowbridge, Frank R.
Black, Merel Game, John Lomer, Frank Popp, Robert Trumbule, Robert B.
Black, Robin S. Gamon, John Lonard, Robert Pregaldin, Charles Tucker, Gordon C.
Blaney, Sean Gardner, Rick Long, R. Pringle, James S. Turner, Billy
Blondeau, Marcel Gilman, Arthur Loope, L.L. Prior, Cathy Ukropina-Crawford, Zvezdana
Bluhm, Wilbur Glenn, S. D. Lucio, Anna Pyne, Milo Underwood, Francis
Boone, D. Goldman, Douglas H. Ludwig, Chris Rabeler, Richard K. Underwood, Malissa
Boufford, David E. Goodman, Darryl MacBryde, Bruce Rader, Linda Valente, Matthew
Bourgeois, David M. Gordon, Ken Magee, Dennis Rawinski, Tom Vincent, Michael
Brandenburg, David Gould, K. R. Magney, David L. Ray, Gary Virag, Gregory
Bregar, John Grafton, Bill Malan, Gerald Read, Robert H. Vizgirdas, Ray
Brouillet, Luc Graham, Shirly A. Mancuso, Michael Rebman, Jon P. Vogel, Johannes
Brown, John Gregory, Linda Mann, William Rector, Barreon Vosen, Harold
Brown, Larry Guala, Gerald Manning, Wayne Rector, Brian G. Voss, Edward
Brown, Melvin Haber, Erich Mansfield, Don Reed, Clyde Vrilakas, Sue
Brown, Paul Martin Haines, Arthur Martine, Chris Reed, Monique Dubrule Wagner, Warren H.
Bryson, Charles Haldeman, Jan Mastrogiuseppe, Joy Reeder, John Wagner, Warren L.
Burkhart, Eric Halliday, Glenn Mayfield, Mark H. Reid, Christopher S. Walasewicz, Steve
Burton, Michael G. Halpern, Alison McAvoy, William Rejmßnek, Marcel Walker, Chuck
Callahan, Frank Halquist, Barre McClendon, Tom Reveal, James L. Ward, Jerome V.
Campbell, Julian Halse, Richard McCook, Lucile M. Reyher, Deanna Ware, Donna
Canne-Hilliker, Judith Hammer, Mike McCoy, Robert Reznicek, Anton Warner, Richard
Caplow, Florence Hansen, Bruce F. McCoy, Roger Rhoads, Ann F. Weakley, Alan S.
Carlson, Paul Hansen, Curtis J. McDaniels, Sidney Rice, Barry Weber, William A.
Carr, William R. Hansen, Jim McDonald, John Robinson, Allan J. Webster, Grady L.
Carter, Louisa Harmon, Paul J. McGregor, Ronald L. Roemer, Brenda Weishaupt, Clara
Carter, Richard Harri, John A. McIlveen, W. D. Roemer, Hans Wellard, Blake
Catling, Paul Harsh, Joseph McLaughlin, Steven P. Rohrer, Joseph Wellford, Alice
Ceska, Adolf Hartman, Ron McMillan, Patrick Roland, Carl Wells, Elizabeth
Chambers, Kenton Harvy, Leroy Meacham, Christopher Rollins, James Werier, David
Charlet, David Haubrich, Gregg Meades, Susan Rollins, Reed C. Westbrooks, Randy
Cholewa, Anita Haynes, Robert R. Mears, James Rondeau, J. Hawkeye White, Deborah
Chumley, Tim Hays, Mike Medley, Max Rosen, David J. White, Peter
Clark, Lynn G. Hegji, Steve Mehrhoff, Leslie J. Sanders, Andrew C. Wieboldt, Thomas
Clemants, Steven Heidel, Bonnie Merkel, Richard Sanders, Julie L. Wieland, Ronald
Clewell, Andrew F. Heil, Ken Meyer, Janice Saulys, Eleanor Wiens, John F.
Clifford, Arnold Heiser, Charles Meyers-Rice, Barry Scheibout, Michael Wiersema, John H.
Cody, William J. Hellquist, Barre Michener, Marty Schimpf, David Wilbur, Robert
Collins, Turner Henderson, James Millman, John Schmid, James Wilczynski, Carolyn
Constance, Lincoln Herbst, Derral Mitchell, Richard S. Schmid, Rudolf Wilder, George
Cook, Mary Beth Hill, Nick Mitrow, G. Schmidt, Greg J. Wilken, Dieter
Cooperrider, Tom S. Hill, Steve Moe, Richard Schneider, Al Williams, John R.
Coxe, Robert Hoagland, Bruce Mohlenbrock, Robert H. Schneider, Betty Williston, Patrick
Cranfill, Ray Hogan, Tim Moldenke, Harold Schneider, Greg Wilson, Barbara
Cronquist, Arthur Homoya, Mike Moore, Jennifer A. Scholz, Hildemar Windham, Michael
Crusca, Fred Horn, Charles N. Moore, Michael Schotz, Al Wofford, B. Eugene
Culpepper, Robyn Horne, Howard E. Morefield, James D. Schreiner, Karen Woods, Michael
Currie, Haynes Horton, Diana Morse, Diana Schuyler, Alfred E. Woolf, Thomas E.
Cusick, Allison W. Hough, Mary Morse, Dorothy V. Semple, John C. Wunderlin, Richard P.
D'Arcy, William G. Howell, J.T. Morse, Larry E. Shell, Paul Yatskievych, George
Damrel, Dixie Hrusa, Fred Morton, Cynthia Sheviak, Charles Yatskievych, Kay
Darst, Melanie R. Hudson, John Moser, Melissa Short, Jack Yenish, Joe
Davis, Cleve Hunt, David Mosyakin, Sergei L. Shory, Rick York, Alan
Davis, Hannibal Hutchens, Robert Mountain, Will Shultz, Leila M. Young, Rodney
Day, Alva G. Hyatt, Philip Murray, David F. Simmons, Roderick H. Zika, Peter
Dempster, Lauramay Imlay, Marc Musselman, Litton Sivinski, Robert Zimmerman, Julie
Diamond, Alvin Ingram, Dianne Naczi, Robert F.C. Smagula, Amy P.  
Dobberpuhl, June Isely, Duane Nation, Fred Smith, Eric L.  
Dorn, Robert D. Jacobs, Don Neal, Joseph C. Smith, J.P.  
Dozier, Ivan Jacono, Colette Neill, Amanda K. Smith, James W.  
Drapalik, Don Jennings, William F. Nekola, Jeff Smith, Latimore  
Duffy, Mike Jensen, Scott Nelson, Burrell B. Smith, Matthew  
Duncan, Tom Johnson, Lewis Nelson, Dean Smith, S. Galen  
Duncan, Wilbur H. Johnston, Barry Nelson, John B. Smith, Welby  
Dunn, David Johnston, June Nesom, Guy L. Snow, Neil W.  


Photographic contributors

Ackerfield, Jennifer R. Fak, Kelly Leonard, Steve Rundel, P.
Alverson, Ed Fay, John Lesica, Peter Rush, Tracy
Amoroso, J. Ferguson, Leslie Lewallen, Charles Rutledge, Paul
Anderberg, Anna Lena Fertig, Walter Lewis, F. Rutman, Susan
Anderberg, Arne Flavon  Lichthardt, Juanita Savela, M.
Anderson, David G. Flemming, G. B. Liston, Aaron Schassberger, Lisa A.
Anderson, Loran C. Fletcher, Reggie Lorain, Chris Schneider, Al (www.swcoloradowildflowers.com)
Anglin, Guy Flethcer, Reggie Lorence, D. H. Schotz, Al
Atwood, Duane Gagné, B. Love, Ken Schotz, Alfred
Baker, Wilson Gagné, W. Lowrey, T. K. Schuller, Reid
Balakrishnan, N. Game, John Lund, Leidulf Schulz, Gary E.
Baldwin, Bruce G. Gamon, John MacLaughlin, Maureen Schwalbe, Cecil
Barrett, Jim Giblin, David Malm, Margaret Seefeldt, Steve
Beaches & Shawes  Gladwin, H. Tim Mancuso, Michael Shaw, Sasha
Bender, Mike Gould, Joyce Manhart, J. R. Sheehan, Mark
Bissett, Nancy Gowans, Edward Mantas, Maria Shelly, Steve
Blake, Jill Gray, Karen Markos, Staci Shirah, Steve
Blanchard, Jon Gray, William (Cyberflora) Marriott, H. Sias, Don
Bleakly, David Greenlee, Jack Marttala, Vern Siddall, Jean
Boothe, B. B. Hammer, Roger L. Mathews, Dan Simard, André
Bourneuf, E. Hanson, Bruce F. Mathews, Dave Sirca, S.
Bradshaw, Jerry Hardie, Linda M. Matson, Steve Sivinski, Robert
Britten, Alan Hardin, E. E. Maur, Bryan Smith, Clifford W.
Brooks, Paula Harris, Jessie M. Mayne, Pete Smith, E. LaVerne
Brown, Bill Harrod, Richy J. McAllister, C. Smith, K.
Brown, Claude Hartman, Ann H. McCarthy, Judy Smith, L.
Brown, Jeff Hays, Mike McCartney, Charles Smith, Matthew
Bruggen, Theodore van Heart, T. McDonald, Charlie Smith, Ruth
Brunsfeld, Robert Hedenas, Lars McGinnes, W. Smith, Welby
Brunsfeld, Steve Heidel, Bonnie McLaughlin, William B. Sole, Mike
Bryan, Dana C. Heil, Ken Medeiros, A. C. Sorrie, Bruce A.
Buckner, Jim Henderson, Douglas Medeiros, Art Spellenberg, Richard
Burkhalter, J. Herbst, D. R. Michener, Martin C. Spence, J.
Burr, D. Herring, Brenda Miller, Anthony Spencer, Marlin
Caicco, Steve Herring, Don Moler, Paul Starr, Forest and Kim (USGS)
Canfield, Joan Hill, Janice Moranz, Ray Steinkamp, Myrna
Caplow, Florence Hillgard, Debra Morden, C. W. Stemmermann, Lani
Caraway, Vickie Hilsenbeck, R. A. Morefield, James D. Stemmermann, R. L.
Carlquist, S. Hipes, Dan Moseley, Robert Stevens, M. L.
Carr, Gerald D. Hobdy, R. Mosley, Robert Stewart, Jon
Carr, M. A. Holland, J. Motley, T. J. Stout, I. Jack
Carr, R. L. Holloway, Vern Müller, Jones Strickler, Dee
Carter, Jack L. Hopkins, Bill Munoz, William Sugii, Nellie
Ceska, Adolf Howard, Mike Murphy, Christopher Suzuki, Kevin
Chafin, Joe Budd/Linda Huppman, Liz Myers, L. Swartz, Linda
Chafin, Linda G. Hutchins, Charles R. Naumann, Tamara Taliapietra-Cherbavaz
Charters, Michael L. Hyatt, Philip E. Neely, Betsy Taylor, Amy
Chauvin, Yvonne Jackson, Dale Nelson, Gil Taylor, Ronald J.
Chimera, C. Jacobi, J. D. Nelson, John Taylor, Walter
Christie, Christopher L. Jacobson, Anders Nelson, Julie Kierstead Tenaglia, Dan
Christman, Steve Jankovsky-Jones, Mable Nesmith, Kathy Tisdale, Edwin
Clemants, Steve Jenkins, C. Nipes, Don Tobe, J. T.
Cole, Nancy Jennings, Bill Obata, J. K. Todd, A. J.
Coleman, Ronald A. Jensen, John Ode, David J. Todd, R.
Coles, Janet Jog, Suneeti O'Kane, Steve Todsen, Tom
Collins, L. T. John, Harold St. Old, Richard (www.xidservices.com) Tonne, Phil
Conant, Sheila Johnson, Ann F. Olsen, M. Toolin, L.J.
Cox, Ann Johnson, Fred Oppenheimer, H. Tuhy, Joel S.
Cox, Robert Johnson, Terry B. Oswald, Vernon H. Urban, Karl
Crins, Bill Johnston, Barry Palis, John Vanderhorst, Jim
Crosby, V. Jones, Loring M. Panjabi, Susan Spackman Vandevender, T.R.
Cuddihy, L. Joy, John Patrick, Grace Vrilakas, Sue
Daehler, C. C. Joyal, Elaine Pavek, Diane Wagner, Warren
Daida, G. Judd, Walter Pearsall, G. A. Walkes, Dick
DeBolt, Ann Kagan, Jimmy Peek, James R. Warshauer, F. R.
DeBruin, Ellen Karl, D. Philips, Barb G. Wasowski, Andy
Degener, O. Keighery, Greg Phillips, Wayne Weldy, Troy
Degener, O. & A. Greenwell  Kemp, Lois Pierce, John Wellner, Chuck
Degener, O. & I.  Kendall, Barry Pierce, Prince Wemersson, B.
Delaney, A. Kershaw, Linda Platt, J. Wenger, D.
Denno, R. F. Kerstetter, Tulli Poole, Jackie Wenier, David
DeSanto, Jerry Kindell, Carolyn Popovich, Steve Wentworth, Jane
Dietrich, Eleanor Klockner, W. Porter, Dallas Whipple, Jennifer
Droege, Mary Knight, Gary Potrick, T.S. Whistler, Art
DuBois, Kristi Knight, Paul Prendusi, Teresa White, D.
Duft, Joe Kores, Paul Randall, Rod White, M.J.D.
Dumaran, Melissa Kosterin, O. Ratzloff, Jim Wiancko, R. Dennis
Dunbar, S. Kral, Robert Rau, Tom Williams, T. Ann
Duncun, W.H. Kratz, Andrew Reed, Daniel W. (www.2bnTheWild.com) Winger, R.
Dunminre, William Kuntz, David Refsdal-Delmatier, Charmaine Wirt, Steve
Eastman, Don Kunze, Linda Reichenbacher, F.W. Wood, Ken
Eastman, Don and Priscilla Kyhos, D. W. Ripple, Jeff Worthington, Richard
Edinger, G. Ladyman, Juanita Ritchie, R. Wright, Carolyn
Egger, Mark Lamoureux, C. H. Rittenhouse, Bruce Wunderlin, Richard
Eldblom, N. Lance, Ron Rodeen Yanuck-Platt, B.
Elliasson, U. Larson, Tammy Roe, Lisa S. Yatskievych, George
Emery Layne, J. N. Rosentreter, Roger Young, Steve M.
Epple, Anne Orth and Lewis E. Lee, Linda Ross, T.L. Jr. Zika, Peter
Evenden, Angie Legler, Ben Roth, Daniela Zimmerman, Dale
Evenson, Bill LeGrande, M. Roth, John E.  

Additional Photographic Sources:

Colorado Natural Areas Program S.M-B SGW University of Georgia Herbarium
Idaho Native Plant Society  Saxifraga Foundation  U.S. Fish & Wildlife Service  US Fish & Wildlife Service Photo File
JRC Seattle FilmWorks University of Florida: Center for Aquatic and Invasive Plants William L. Lunk in P.D. Strausbaugh and E.L. Core. 1978. Flora of West Virginia
Oregon Biodiversity Information Center      


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Last updated October 11, 2011.